TY - RPRT TI - Coordinated immune, chloroplast and chemical defences underpin multilayered resistance to barley yellow dwarf virus and its aphid vector associated with the Hordeum bulbosum-derived Ryd4 introgression in barley AU - Simon, A. AU - Jayaweera, D. AU - Qonaah, I. A. AU - Verburg, D. AU - Clarke, F. AU - Kim, D.-H. AU - Urquhart, B. AU - Melichar, J. AU - Giles, T. AU - Bruce, T. AU - Ray, R. V. PY - 2026 DO - 10.64898/2026.07.28.741184 UR - https://www.biorxiv.org/content/10.64898/2026.07.28.741184v1 ID - 10.64898/2026.07.28.741184 ER -